north-west of the Cerro Leon, Alto Paraguay, Paraguay, 164 m
north-west of the Cerro Leon, Alto Paraguay, Paraguay, 164 m
east of Charagua towards Yapiroa, Santa Cruz, Bolivia, 606 m
guanchinense v. mirandense, STO 88
Cuesta Miranda, Las Higueras, La Rioja, Arg.
marekiorum subsp. sanjoseanum, VoS 1876
east of Santa Cruz, west of Rio Grande, Santa Cruz, Bolivia, 307 m
north-east of Fortin Ravello, Santa Cruz, Bolivia, 291 m